Board Thread:Spoilers!/@comment-6175354-20140420035659/@comment-6175354-20140424173646

DZAV21 wrote: I think giving up what they loved most was symbolic like when Regina gave up Henry in the midseason final,

If it was just a symbolic sacrifice, vs. crushing an actual heart, then the caster was Hook. He is the only person in town with memories, and the Jolly Roger is missing.

Though, I think the creators said, that someone's heart was sacrified.